Three papers cited by your link:
M.J.C. Crabbe of the United Kingdoms University of Bedfordshire
concluded Despite the multiple influences on the reef sites over the study period, the size classes of the corals studied showed resilience to change. We suspected this all along the coral reefs have been around for 100s of millions of years! He states What is apparent from this study is that despite the chronic and acute disturbances between 2002 and 2008, demographic studies indicate good levels of coral resilience on the fringing reefs around Discovery Bay in Jamaica. Crabbe warns that Unfortunately, previously successful efforts to engage the local fisherman in controlling catches around Discovery Bay have not been maintained, and it may be that the development of a Discovery Bay Marine Park is the only solution. We get the message dont blame global warming, blame the local fishermen!
Study limited to localized area where the author believes that overfishing is a factor. Temperature infulences have not been ruled out.
Mumby and Harborne
Because the Bahamas was severely disturbed by the 1998 coral bleaching event, and later by hurricane Frances in the summer of 2004, coral cover was low at the beginning of the study, averaging only 7% at reserve and non-reserve sites. Corals have been around for eons, they have survived periods much hotter than anything experienced today, they have survived massive El Niño events, and as seen in their study area, the corals can be severely damaged by hurricanes. Delicate corals would have never made it robust corals would win in the world of natural selection.
While this might be true, temperature change was more gradual both before and after the higher temperature change. What is happening now is more rapid. True some coral species will be able to survive, but this will mean a change in the coral reef fauna and flora. However as temperatures continue to raise there can be more drastic die off.
Adjeroud et al. studied the Tiahura Outer Reef Sector
Their conclusions include the statement In addition, our results reveal that corals can recover rapidly following a dramatic decline. Such decadal-scale recovery of coral cover has been documented at some locations, but our results are novel in demonstrating rapid recovery against a backdrop of ongoing, high frequency, and large-scale disturbances.
The three papers cited in your article mentions the resiliency of coral reefs. The paper I cited does not dispute this. What it does point out that coral reefs are being impacted faster than they can recover. Corals are very sensitive to temperatures, they live in regions where temperatures are more stable. While there could be multiple factors involved, as the paper I cited acknowledges, temperatures have the greatest impact. Sea temperatures of 2010 have exceeded 2005's the last time this occurred.Scientists: Caribbean coral die-off may be worst ever, Southeast Asia and Indian Ocean bleaching may prove to be the worst such event known to science.
Ive never seen bleaching like [it] in Panama, said Nancy Knowlton, a coral biologist at the Smithsonian Tropical Research Institute in Panama who has been studying the local flora for 25 years. She and colleague Hector Guzman have seen massive reefs die in recent weeks in the enclosed lagoon of Bocas del Toro in Panama after becoming coated with giant sheets of slime, the remains of dead microorganisms. This is NOT a normal condition on reefs, even bleached reefs. Where last year there were healthy corals, this year there was only gray ooze, she wrote in an e-mail.
A number of factors besides water temperature can cause coral bleaching and die-offs, including pollution and storms. But temperature is the number-one culprit in such a massive die-off, says Eakin. The warmest 12-month period in the NASA temperature record ended this summer; June through August was the fourth-warmest such period in the record. The extent of the devastation across the Caribbean will become clear in the coming months as biologists measure the deaths.